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Google Scholar page.

Underlined names denote undergraduate coauthors.

Book Chapters and Popular Press

  • Keiser CN. Collective behavior and parasite transmission. 2022. In: Ezenwa VO, Altizer S, Hall RJ, editors. Animal Behavior and Parasitism. Oxford University Press, 2022. [PDF]

  • Bahlai CA, Bartlett LJ, Burgio KR, Fournier AMV, Keiser CN, Poisot T, and Whitney KW. 2018. Open science isn’t always open to all scientists. American Scientist.107: 78-82. [PDF]

  • Keiser CN, Lichtenstein JLL, Wright CM, Dittrich-Reed D, Chism GT, Jonathan N. Pruitt. 2016. Personality and behavioral syndromes in insects and spiders. In Insect Behavior: From Mechanisms to Ecological and Evolutionary Consequences. Oxford University Press. [Book chapter]


  • Durkin ES, Cassidy ST, Leyva A, and Keiser CN. Population differences
    in the aggregation and collective foraging behavior of fra
    gmented social spider
    Ethology. 129: 224-231. [PDF] [data]


  • Cassidy ST, Kianna Saadatmand, Yinlu Zhu, Alice Gau, Chelsey Gerena,
    Keiser CN. The effect of social parasite exposure on an
    t colony defenses. 

    Journal of Insect Behavior. 36: 52-58. [PDF] [data]

  • Nyffeler M, Machado G, Hartmann AG and Keiser CN. Vegetarian arachnids:
    a review on fungi and fruit consumption in harvestmen and spiders
    (Opiliones and Araneae).
    Journal of Arachnology. 51: 1-18. [PDF]


  • Richardson EA, Buttrick DJ, Shablin SA, Jabot B, Taylor CE, Martin EM, and Keiser CN. 2022. Exploring the effects of pathogen infection on tick behavior from individuals to populations. Animal Behaviour. 186: 85-92. [PDF] [data] [code]

  • Richardson, EA, Taylor CE, Jabot B, Martin E, Keiser CN. 2021. The effects of habitat type and pathogen infection on tick host-seeking behavior. Parasitology. [PDF] [data]



  • Durkin ED, Cassidy ST, Gilbert R, Richardson, EA, Roth AM, Shablin S, Keiser CN. 2021. Parasites of spiders: Their impact on host behavior and ecology. Journal of Arachnology. 49(3): 281-298. [PDF]

  • Cassidy SC, Chapa J, Tran T, Dolezal N, Gerena C, Johnson G, Leyva A, Stein S, Wright CM, and Keiser CN. 2021. Immune defenses across levels of biological organization: Individual and social immunity in acorn ants. Accepted/In press. Animal Behaviour. 179: 73-81. [PDF] [data]

  • Roth AM, Keiser CN, Williams JB, Gee, JM. 2021. Prevalence and intensity of avian malaria in a quail hybrid zone. Ecology and Evolution. 11: 8123-8135. [PDF] [data]

  • Roth AM, Gee J, and Keiser CN. 2021. Autumn copulatory behavior in California Quail: Observations and potential functions. Western Birds. 52: 112-123. [PDF]



  • Durkin ES, Roth AR, Keiser CN. 2020. Parasitic personalities: Consistent individual differences in behavior in a facultatively parasitic mite. Journal of Insect Behavior. [PDF] [data].

  • ​Keiser CN, Rudolf VHW, Luksik MC, and Saltz JB. Sex-differences in disease avoidance behavior vary across modes of pathogen exposure. Ethology. 126: 304-312. [PDF] [data]

  • Wright CM, Pinter-Wolman N, Geary S, Pruitt JN, Keiser CN. Spatial proximity and prey vibratory cues influence collective hunting in social spiders. Israel Journal of Ecology and Evolution. 66: 26-31.



  • Ul-Hasan S, Volker Herzig, Rachelle M Adams, Eduardo Rodríguez-Román, Steven A Trim, Adam R Reitzel, Sterghios A Moschos, Clarissa J Nobile, Keiser CN; Daniel Petras, Erin E Stiers, and Colston TJ. Exploring the venom ecosystem: The emerging field of venom-microbiomics and the Initiative for Venom Associated Microbes and Parasites. Toxicon. 4: 100016. [PDF]

  • Keiser CN, Hammer TJ, and Pruitt JN. Social spider webs harbor largely consistent bacterial communities across broad spatial scales.
    Biology Letters. 15: 20190436. [PDF] [data]


  • Keiser CN, Wantman T, Rebollar EA, and Harris RN. Tadpole body size and behavior alter the social acquisition of a defensive bacterial symbiont. Royal Society Open Science. 6:191080. [PDF] [data]

  • Spicer ME, Pruitt JN, and Keiser CN. Spiders, microbes, and sex: Bacterial exposure on copulatory organs alters mating behaviour in funnel-web spiders. Ethology. 125: 677-685. [PDF] [data]



  • Pruitt JN, Keiser CN, Banka BT, Leidle JS, Brooks AJ, Schmitt RJ, Holbrook SJ. 2018. Collective aggressiveness of animal groups enhances reestablishment of a foundation species. Behavioral Ecology. 29: 1216-1224.

  • Keiser CN, Rudolf VHW, Sartain E, Every E, and Saltz JB. 2018. Social context alters host behavior and infection risk. Behavioral Ecology. 29: 869-875. [PDF] [data]

  • Keiser CN, Ingley SJ, Toscano BJ, Scharf I, and Pruitt JN. 2018. Habitat complexity dampens selection on prey activity level. Ethology. 124: 25-32. [PDF] [data]

  • Keiser CN, Pinter-Wollman N, Ziemba MJ, Kothamasu KS, and Pruitt JN. 2018. The primary case is not enough: exploring trait variation among individuals and social groups in bacterial transmission dynamics. Journal of Animal Ecology87: 369-378. [PDF] [data]

  • Keiser CN, Vodjvodic S, Butler IO, Sartain E, Rudolf VHW, and Saltz JB. 2018. Queen presence alters the relationship between collective behavior and disease susceptibility in ant colonies. Journal of Animal Ecology. 87: 379-387. [PDF] [data]


  • Parks OB, Kothamasu K, Ziemba M,… Keiser CN. 2017. Increased cuticular bacterial load alters aggressiveness in a funnel-weaving spider. Current Zoology. zox064. [PDF] [data]


  • Keiser CN, Howell KA, Pinter-Wollman N, and Pruitt JN. 2016. Personality composition alters the transmission of cuticular bacteria in social groups. Biology Letters. 12: 20160297. [PDF] [data]

  • Pinter-Wollman N, Keiser CN, Wollman R, and Pruitt JN. 2016. The effect of keystone individuals on tradeoffs between collective outcomes can be mediated through interaction rules or behavioral persistence. The American Naturalist. 188: 240-252. [PDF] [data]

    • CORRECTION: Pinter-Wollman N, Keiser CN, Wollman R. 2021. The American Naturalist [link]

  • Keiser CN, Pinter-Wollman N, Augustine DA, Ziemba MJ, Hao L, Lawrence JG, and Pruitt JN. 2016. Individual differences in boldness influence patterns of social interactions and the transmission of cuticular bacteria among group-mates. Proceedings of the Royal Society B: Biological Sciences. 283: 20160457. [PDF] [data]

  • Wright CM, Keiser CN, and Pruitt JN. 2016. Colony personality composition alters colony-level plasticity and magnitude of defensive behaviour in a social spider. Animal Behaviour. 115: 175–183. [PDF]

  • Keiser CN, Wright CM, and Pruitt, JN. 2016. Increased bacterial load on keystone individuals alters group collective behaviour. Animal Behaviour. 114: 211-218.  [PDF]

  • Keiser CN, Shearer TA, DeMarco AE, Brittingham HA, Knutson KA, Kuo C, Zhao K, and Pruitt JN. 2016. Cuticular bacteria prove lethal to social spiders in mixed but not monoculture inoculations. Current Zoology. [PDF]


  • Modlmeier AP, Laskowski KL, Brittingham HA, Coleman A, Knutson KA, Kuo C, McGuirk M, Zhao KC, Keiser CN, Pruitt JN. 2015. Adult presence augments juvenile collective foraging in social spiders. Animal Behaviour. 109: 9-14. [PDF]

  • Keiser CN, Wright CM, and Pruitt, J.N. 2015. Warring arthropod societies: Colony apparency and group size jointly dictate the survivorship of social spiders in a predator-dense habitat. Behavioural Processes. 119: 14-21. [PDF]

  • Keiser CN, DeMarco AE, Shearer TA, Robertson JA, and Pruitt JN. 2015. Putative microbial defenses in a social spider: Immune variation and antibacterial properties of colony silk. Journal of Arachnology. 43: 394-399. [PDF]

  • Keiser CN, Slyder JB, Carson WP, and Pruitt JN. 2015. Individual differences in predators but not producers mediate the magnitude of a trophic cascade. Arthropod-Plant Interactions. 9: 225-232. [PDF] [data]

  • Modlmeier AP, Keiser CN, Wright CM, Lichtenstein JL, and Pruitt JN. 2015. Integrating animal personality into insect population ecology. Current Opinions in Insect Science. 9: 77-85. [PDF]

  • Keiser CN, Wright CM, Singh N, DeShane JA, Modlmeier AP, and Pruitt JN. 2015. Cross-fostering by foreign conspecific queens and slavemaking workers influences individual and colony-level personality. Behavioral Ecology and Sociobiology. 69: 395-405.

  • Keiser CN and Mondor EB. 2015. Cues of predation risk induce instar- and genotype-specific changes in pea aphid colony spatial structure. Ethology. 121: 144-151.


  • Keiser CN and Pruitt JN. 2014. Personality composition is more important than group size in determining collective foraging behaviour in the wild. Proceedings of the Royal Society B: Biological Sciences. 281: 20141424. [PDF] [data]

    • CORRECTION [link]

  • Modlmeier AP, Keiser CN, Shearer TA, and Pruitt JN 2014. Species-specific influence of group composition on collective behaviors. Behavioral Ecology and Sociobiology. 68: 1929-1937. [PDF]

  • Keiser CN and Pruitt JN 2014. Submersion tolerance in a lakeshore population of Pardosa lapidicina (Araneae: Lycosidae). Journal of Arachnology. 42:192-194. [PDF]

  • Keiser CN, Modlmeier AP, Singh N, Jones DK, and Pruitt JN. 2014. Exploring how a shift in the physical environment shapes individual and group behavior across two social contexts. Ethology. 120: 825-833. [PDF] [data]

  • Keiser CN, Jones DK, Modlmeier AP, and Pruitt JN. 2014. Exploring the effects of individual traits and within-colony variation on task differentiation and collective behavior in a desert social spider. Behavioral Ecology and Sociobiology. 68: 839-850. [PDF] [data]

  • Modlmeier AP, Keiser CN, Watters JV, Sih A, and Pruitt JN. 2014. The keystone individual concept: an ecological and evolutionary overview. Animal Behaviour. 89: 53-62. [PDF]

2013 and earlier

  • Keiser CN, Sheeks LE, and Mondor EB. 2013. The effect of microhabitat feeding site selection on aphid foraging and predation risk. Arthropod-Plant Interactions. 6: 633-642.

  • Pruitt JN and Keiser CN. 2013. Debates: The aggressive spillover hypothesis: existing ailments and putative remedies. Ethology. 119: 807-810.

  • Sweeney K, Cusack B, Armagost F, O'Brian T, Keiser CN, and Pruitt JN. 2013. Predator and prey activity levels jointly influence the outcome of long-term foraging bouts. Behavioral Ecology. 24: 1205-1210.

    • CORRECTION: [link]

  • Keiser CN and Mondor EB. 2013. Transgenerational behavioral plasticity in a parthenogenetic insect in response to increased predation risk. Journal of Insect Behavior. 26: 603-613.

  • Mondor EB, Keiser CN, Pendarvis D, and Vaughn MN. 2013. Broad bean cultivars increase extrafloral nectary numbers, but not extrafloral nectar, in response to leaf damage. Open Journal of Ecology. 3: 59-65.

  • Hoefler CD, Keiser CN, and Rypstra AL. 2010. Sex differences in early instar behavior in Pholcus phalangioides (Araneae, Pholcidae). Journal of Arachnology. 38: 581-583.

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